Hannier Pulido, Gonzalo Andrade, Carlos Peña, Gerardo Lamas. 2011. Two new taxa of Euptychia Hübner, 1818 (Lepidoptera: Nymphalidae: Satyrinae) from the Andes of Colombia and Peru. Zootaxa : 43-51.

Carlos Peña, Sören Nylin, Niklas Wahlberg. 2011. The radiation of Satyrini butterflies (Nymphalidae: Satyrinae) - a challenge for phylogenetic methods. Zoological Journal of the Linnean Society 161: 64-87.

Carlos Peña, Sören Nylin, A.V.L. Freitas, Niklas Wahlberg. 2010. Biogeographic history of the subtribe Euptychiina (Lepidoptera, Nymphalidae, Satyrinae). Zoologica Scripta 39: 243-258.

Ullasa Kodandaramaiah, Carlos Peña, Michael F. Braby, Roger Grund, Chris J. Müller, Sören Nylin, Niklas Wahlberg. 2010. Phylogenetics of Coenonymphina (Nymphalidae Satyrinae) and the problem of rooting rapid radiations. Molecular Phylogenetics and Evolution 54: 386-394.

Niklas Wahlberg, Julien Leneveu, Ullasa Kodandaramaiah, Carlos Peña, Sören Nylin, André V L Freitas, Andrew V Z Brower. 2009. Nymphalid butterflies diversify following near demise at the Cretaceous/Tertiary boundary. Proceedings of the Royal Society of London, Series B, Biological Sciences 276: 4295-4302.

Carlos Peña, Niklas Wahlberg. 2008. Prehistorical climate change increased diversification of a group of butterflies. Biology Letters 4: 274-278.

Carlos Peña, Niklas Wahlberg, Elisabet Weingartner, Ullasa Kodandaramaiah, Sören Nylin. 2006. Higher level phylogeny of Satyrinae butterflies (Lepidoptera: Nymphalidae) based on DNA sequence data. Molecular Phylogenetics and Evolution 40: 29-49.

André Freitas, Carlos Peña. 2006. Description of genus Guaianaza for "Euptychia" pronophila (Lepidoptera: Nymphalidae: Satyrinae) with a description of the immature stages. Zootaxa 1163: 49-59.

Carlos Peña, Gerardo Lamas. 2005. Revision of the butterfly genus Forsterinaria Gray, 1973 (Lepidoptera: Nymphalidae, Satyrinae). Revista peruana de Biología 12: 5-48.

Other academic

Carlos Peña. 2009. Evolutionary history of the butterfly subfamily Satyrinae (Lepidoptera: Nymphalidae). Doctoral thesis. Department of Zoology, Stockholm University.

Other (popular science, discussion, etc.)

Niklas Wahlberg, Carlos Peña. 2007. A Dominican Republic sojourn. News of the Lepidopterists' Society 49: 12-15, 37.